Mackinnon Memorial Hospital

Introduction.Following on from work on the European bryophyte Red List, the taxonomically and nomenclaturally updated spreadsheets used for that project have been expanded into a new checklist for the bryophytes of Europe.Methods.A steering group of ten European bryologists was convened, and over the course of a year, the spreadsheets were compared with previous European checklists, and all changes noted.Recent literature was searched extensively.A taxonomic system was agreed, and the advice and expertise of many European bryologists sought.Key results.A new European checklist of bryophytes, comprising hornworts, liverworts and mosses, is presented.Fifteen new combinations are proposed.Conclusions.This checklist provides a snapshot of the current European bryophyte flora in 2019.It will already be out-of-date on publication, and further research, particularly molecular work, can be expected to result in many more changes over the next few years.

Orkney was a major cultural center during the Neolithic, 3800 to 2500 BC. Farming flourished, permanent stone settlements and chambered tombs were constructed, and long-range contacts were sustained. From ∼3200 BC, the number, density, and extravagance of settlements increased, and new ceremonial monuments and ceramic styles, possibly originating in Orkney, spread across Britain and Ireland. By ∼2800 BC, this phenomenon was waning, although Neolithic traditions persisted to at least 2500 BC. Unlike elsewhere in Britain, there is little material evidence to suggest a Beaker presence, suggesting that Orkney may have developed along an insular trajectory during the second millennium BC. We tested this by comparing new genomic evidence from 22 Bronze Age and 3 Iron Age burials in northwest Orkney with Neolithic burials from across the archipelago. We identified signals of inward migration on a scale unsuspected from the archaeological record: As elsewhere in Bronze Age Britain, much of the population displayed significant genome-wide ancestry deriving ultimately from the Pontic-Caspian Steppe. However, uniquely in northern and central Europe, most of the male lineages were inherited from the local Neolithic. This suggests that some male descendants of Neolithic Orkney may have remained distinct well into the Bronze Age, although there are signs that this had dwindled by the Iron Age. Furthermore, although the majority of mitochondrial DNA lineages evidently arrived afresh with the Bronze Age, we also find evidence for continuity in the female line of descent from Mesolithic Britain into the Bronze Age and even to the present day.

Abstract Bryophytes are a diverse group of organisms with unique properties, yet they are severely underrepresented in plant trait databases. Building on the recently published European Red List of bryophytes and previous trait compilations, we present the Bryophytes of Europe Traits (BET) data set, including biological traits such as those related to life history, growth habit, sexual and vegetative reproduction; ecological traits such as indicator values, substrate and habitat; and bioclimatic variables based on the species' European range. The data set includes values for 65 traits and 25 bioclimatic variables, containing more than 135,000 trait values with a completeness of 82.7% on average. The data set will enable future studies in bryophyte biology, ecology and conservation, and may help to answer fundamental questions in bryology.

1. Asterella lindenbergiana (Corda ex Nees) ArnellContributors: E. A. Borovichev and V. A. BakalinRussian Federation (the Russian Far East): Magadan Province, Magadan City vicinity, Gertnera Bay, a...

1. Anoectangium stracheyanum Mitt.Contributors: P. Sollman and A. Schäfer-VerwimpCanada: British Columbia, Vancouver Island, Barclay Sound, 49°04'N, 125°09'W, damp cliff near cascade near head of E...

The Middle Jurassic is a largely mysterious interval in dinosaur evolution, as few fossils of this age are known worldwide. In recent years, the Isle of Skye has yielded a substantial record of trackways, and a more limited inventory of body fossils, that indicate a diverse fauna of Middle Jurassic dinosaurs living in and around lagoons and deltas. Comparatively little is known about the predators in these faunas (particularly theropod dinosaurs), as their fossils are among the rarest discoveries. We here report two new isolated theropod teeth, from the Valtos Sandstone Formation and Lealt Shale Formation of Skye, which we visualized and measured using high-resolution x-ray computed microtomographic scanning (µCT) and identified via statistical and phylogenetic analyses of a large comparative dental dataset. We argue that these teeth most likely represent at least two theropod species – one small-bodied and the other large-bodied – which likely belonged to one or several clades of basal avetheropods (ceratosaurs, megalosauroids, or allosauroids). These groups, which were diversifying during the Middle Jurassic and would become dominant in the Late Jurassic, filled various niches in the food chain of Skye, probably both on land and in the lagoons. Supplementary material : Character lists, datasets, and measurements are available at https://doi.org/10.6084/m9.figshare.c.4452533

1. Afoninia dahurica Ignatova, Goffinet & FedosovContributors. D. Ya. Tubanova and O. D. DugarovaRussia. Republic of Buryatia: Zakamenskiy District, Dzhidinskiy Ridge, about 10 km to SSE of Mikhayl...

1. Acrobolbus ciliatus Mitt.Contributors: D. Gupta, K. K. Rawat, V. Sahu and A. K. AsthanaIndia: Uttarakhand, Uttarkashi, Govind Wildlife Sanctuary, between Badang and Daldhar, Vijay top, 31°08.86...

Atoposaurids were a clade of semiaquatic crocodyliforms known from the Late Jurassic to the latest Cretaceous. Tentative remains from Europe, Morocco, and Madagascar may extend their range into the Middle Jurassic. Here we report the first unambiguous Middle Jurassic (late Bajocian-Bathonian) atoposaurid: an anterior dentary from the Isle of Skye, Scotland, UK. A comprehensive review of atoposaurid specimens demonstrates that this dentary can be referred to Theriosuchus based on several derived characters, and differs from the five previously recognized species within this genus. Despite several diagnostic features, we conservatively refer it to Theriosuchus sp., pending the discovery of more complete material. As the oldest known definitively diagnostic atoposaurid, this discovery indicates that the oldest members of this group were small-bodied, had heterodont dentition, and were most likely widespread components of European faunas. Our review of mandibular and dental features in atoposaurids not only allows us to present a revised diagnosis of Theriosuchus, but also reveals a great amount of variability within this genus, and indicates that there are currently five valid species that can be differentiated by unique combinations of dental characteristics. This variability can be included in future broad-scale cladistics analyses of atoposaurids and closely related crocodyliforms, which promise to help untangle the complicated taxonomy and evolutionary history of Atoposauridae.

Synopsis The first in situ dinosaur tracks from Scotland were discovered at the top of the Duntulm Formation (Bathonian, Jurassic) near to Staffin in northeastern Skye. Fifteen individual tridactyl footprints were recorded of which two pairs appear to have been part of the same trackway. The footprints are preserved as natural moulds on a mud-cracked sandstone surface. The individual track sizes range from about 30 cm to over 50 cm in length with narrow to broad digits suggestive of having been made by a medium to large bipedal dinosaur.

The IUCN Red List is recognised as a robust system for assessing the risk of extinction to organisms, but there are difficulties in applying the criteria to bryophytes and other clonal and colonial organisms. Three critical terms are addressed – generation length, mature individual and severe fragmentation – and definitions given in order to facilitate the use of the IUCN Red List criteria for bryophytes. These recommendations provide a pragmatic and effective way of using the IUCN Red List process for bryophytes and may have a wider application to other clonal organisms.

Tracks of a juvenile theropod dinosaur with footprint lengths of between 2 and 9 cm as well as adults of the same ichnospecies with footprints of about 15–25 cm in length were found in the Bathonian (Middle Jurassic) Kilmaluag Formation of Score Bay, northwestern Trotternish Peninsula, Isle of Skye, Scotland, UK. Two footprint sizes occur together on the same bedding plane in the central portion of Score Bay, both in situ and on loose blocks. Another horizon containing footprints above this was also identified. The footprints from the lowest horizon were produced in a desiccated silty mud that was covered with sand. A close association of both adults and juveniles with similar travel direction indicated by the footprints may suggest post-hatching care in theropod dinosaurs. Other footprints, produced on a rippled sandy substrate, have been found on the slightly higher bedding plane at this locality. Loose blocks found 130 m to the northeast in the central part of Score Bay have not been correlated with any in situ sediments, but were preserved in a similar manner to those from the higher bedding plane. These tracks represent the youngest dinosaur remains yet found in Scotland.

Introduction. Many taxa have been added to the bryophyte flora of Britain and Ireland since the publication of the previous checklist in 2008. This has coincided with numerous taxonomic and nomenclatural changes in the European flora, especially as a consequence of molecular sequencing. With the publication of a new European checklist in 2020 it is now appropriate to provide an updated and annotated list for Britain and Ireland.Methods. A working group was convened to consider and reach a consensus on taxonomic and nomenclatural matters arising from the European checklist. Special consideration was given to the citation of the authors of names. Taxa reported as new to Britain and Ireland since the previous checklist in 2008 were incorporated.Key results. A new checklist of bryophytes (hornworts, liverworts and mosses) is presented. Based on the taxonomy adopted in the checklist, the bryophyte flora of Britain and Ireland comprises four hornworts (Anthocerotophyta), 299 liverworts (Marchantiophyta), and 795 mosses (Bryophyta) at species rank, with 42 additional infraspecific taxa.

The Middle Jurassic was a dynamic interval in dinosaur evolution, but the dinosaur fossil record from this time is extremely poor throughout the world. The Isle of Skye (Scotland) preserves marginal marine and terrestrial deposits of Middle Jurassic age, which have yielded sparse bones, teeth, footprints and small segments of trackways belonging to dinosaurs. We report the discovery of the most extensive dinosaur fossil site yet known in Scotland: a coastal outcrop of the Duntulm Formation (Bathonian) at Cairidh Ghlumaig, Skye that preserves numerous trackways of sauropod dinosaurs in multiple layers deposited in a lagoonal system. We present an initial description of these tracks and identify them as most likely belonging to a primitive, non-neosauropod species that retained a large claw on manual digit I and produced narrow-gauge trackways. They provide additional evidence that basal sauropods persisted deep into the Middle Jurassic, a time when the earliest members of larger and more derived sauropod lineages were radiating. The new Skye tracks document multiple generations of sauropods living within the lagoonal environments of Jurassic Scotland, and along with other tracks found over the past two decades, suggest that sauropods may have frequented such environments, contrary to their image as land-bound behemoths.

Lymphocyte migration into fresh and preserved peripheral nerve allografts was assessed to determine the effects of preservation time, preservation temperature, and graft harvest technique on the immunologic response to the peripheral nerve allograft. Peroneal nerve was harvested from either live or cadaveric (tissue) donors and stored as 1.5-cm segments at 5 degrees C or 37 degrees C for 1, 3, 5, or 7 days. Each of nine outbred ewes then received multiple segments of peroneal autograft, fresh allograft, and preserved nerve allograft implants. Lymphocyte migration was studied 7 days after implantation by intravenous injection of autologous 111In-labeled lymphocytes and quantified by gamma counter. Lymphocyte migration into fresh allografts (7212 +/- 1575) increased an average of 4.1 times over fresh autograft tissue (1758 +/- 421; p < 0.05). Short-term preservation (24 hours) at both temperatures enhanced lymphocyte migration into pretreated allograft tissue (12684 +/- 2575 at 5 degrees C, 8751 +/- 1577 at 37 degrees C) as compared with fresh allograft (7212 +/- 1575). Conversely, 7 days of pretreatment at both 5 degrees C (3586 +/- 1421) and 37 degrees C (1570 +/- 414) resulted in migration values not significantly different from autograft. No statistically significant difference was seen between grafts harvested from live (5710 +/- 1651) versus cadaveric (tissue) donors (4013 +/- 832) after 5 days of cold preservation.

1. Acroporium praelongum Dixon var. aciphylloides B.C.TanContributors. N. Norhazrina and N. SyazwanaPeninsular Malaysia. Perak, Gunung Korbu, along the way from Korbu Raya to puncak Korbu, 4°40'59....

Contributor. R. OchyraMadagascar. Diana Region, Tsaratanana Massif, 14°01′S, 48°56′W, 1200–2400 m a.s.l., April 1924, leg. Perrier de la Bathe s.n. (H-Brotherus, JE-Herzog, PC-P.de la Varde 0707821...

Globally there is increasing concern about the decline in the health of oak Quercus trees. The impact of a decline in oak trees on associated biodiversity, species that utilize oak trees, is unknown. Here we collate a database of all known birds, bryophytes, fungi, invertebrates, lichens and mammals that use oak (Quercus petraea and Q. robur) in the UK. In total 2300 species are listed in the database. For each species we provide a level of association with oak, ranging from obligate (only found on oak) to cosmopolitan (found on a wide range of other tree species). Data on the ecology of each oak associated species was collated: part of tree used, use made of tree (feeding, roosting, breeding), age of tree, woodland type, tree form (coppice, pollarded, or natural growth form) and season when the tree was used. Data on use or otherwise by each of the 2300 species of 30 other tree species was also collated. A complete list of data sources is provided. For further insights into how this data can be used see Collapsing foundations: The ecology of the British oak, implications of its decline and mitigation options [1]. Data can be found at EIDC https://doi.org/10.5285/22b3d41e-7c35-4c51-9e55-0f47bb845202.

ABSTRACT The Middle Jurassic is a poorly sampled time interval for non-pelagic neosuchian crocodyliforms, which obscures our understanding of the origin and early evolution of major clades. Here we report a lower jaw from the Middle Jurassic (Bathonian) Duntulm Formation of the Isle of Skye, Scotland, UK, which consists of an isolated and incomplete left dentary and part of the splenial. Morphologically, the Skye specimen closely resembles the Cretaceous neosuchians Pachycheilosuchus and Pietraroiasuchus , in having a proportionally short mandibular symphysis, shallow dentary alveoli and inferred weakly heterodont dentition. It differs from other crocodyliforms in that the Meckelian canal is dorsoventrally expanded posterior to the mandibular symphysis and drastically constricted at the 7th alveolus. The new specimen, together with the presence of Theriosuchus sp. from the Valtos Formation and indeterminate neosuchians from the Kilmaluag Formation, indicates the presence of a previously unrecognised, diverse crocodyliform fauna in the Middle Jurassic of Skye, and Europe more generally. Small-bodied neosuchians were present, and ecologically and taxonomically diverse, in nearshore environments in the Middle Jurassic of the UK.

1. Aloina rigida (Hedw.) Limpr.Contributors: O. Yu. Pisarenko, V. E. Fedosov and V. A. BakalinRussia: Primorsky Territory, Dalnegorsky District, vicinity of Dalnegorsk Town, NE-facing steep slope o...